Beetle
Families and Subfamilies of the World
ORDER COLEOPTERA Linnaeus, 1758
Suborder ARCHOSTEMMATA Kolbe, 1908
1. OMMATIDAE Sharp & Muir, 1912
(= Ommadidae; incl. Tetraphaleridae)
TETRAPHALERINAE
Crowson, 1962
OMMATINAE Sharp & Muir, 1912
2. CROWSONELLIDAE Iablokoff-Khnzorian,
1983
3. MICROMALTHIDAE Barber,
1913 - Telephone-pole beetles
4. CUPEDIDAE Laporte, 1836 - Reticulated beetles
(= Cupesidae)
Suborder MYXOPHAGA Crowson, 1955
5. LEPICERIDAE Hinton, 1936
(= Cyathoceridae)
6. TORRIDINCOLIDAE Steffan, 1964
7. HYDROSCAPHIDAE LeConte, 1874 - Skiff beetles
8. MICROSPORIDAE Crotch, 1873 - Minute bog beetles
(= Sphaeriidae)
Suborder ADEPHAGA Schellenberg, 1806
9. GYRINIDAE Latreille, 1810 - Whirligig beetles
SPANGLEROGYRINAE Folkerts, 1879
GYRININAE Latreille, 1810
10. HALIPIDAE Aubé, 1836 - Crawling water beetles
11. TRACHYPACHIDAE C. G. Thomson, 1857
12. NOTERIDAE C. G. Thomson, 1860 - Burrowing water beetles
(incl. Phreatodytidae)
PHREATODYTINAE Uéno 1957
NOTERINAE C. G. Thomson, 1860
13. AMPHIZOIDAE LeConte, 1853 - Trout-stream beetles
14. HYGROBIIDAE Régimbart, 1878
(= Pelobiidae)
15. DYTISCIDAE Leach, 1815 - Predacious diving beetles
COPELATINAE Van den Branden,
1885
LACCOPHILINAE Gistel, 1856
HYDROPHORINAE Aubé, 1836
COLYMBETINAE Erichson, 1837
DYTISCINAE Leach, 1815
AUBEHYDRINAE Guignot, 1842
16. RHYSODIDAE Laporte, 1840 - Wrinkled bark beetles
17. CARABIDAE Latreille, 1802 - Ground and tiger beetles
(incl. Cicindelidae, Paussidae, etc.)
PAUSSINAE Latreille, 1807
GEHRINGIINAE Darlington, 1933
OMOPHRONINAE Bonelli, 1810
CARABINAE Latreiile, 1802
CICINDELINAE Latreille, 1802
HILETINAE Schiodte, 1847
LORICERINAE Bonelli, 1810
ELAPHRINAE Latreille, 1802
MIGADOPINAE Chaudoir, 1861
SIAGONINAE Bonelli, 1810
SCARITINAE Bonelli, 1810
TRECHINAE Bonelli, 1810
HARPALINAE Bonelli, 1810
incertae sedis
PSEUDOMORPHINAE Bonelli, 1810
BRACHININAE Bonelli, 1810
Suborder POLYPHAGA Emery, 1886
Series STAPHYLINIFORMIA Lameere, 1900
Superfamily HYDROPHILOIDEA
Latreille, 1802 (incl. Histeroidea)
18. HYDROPHILIDAE Latreille, 1802 - Water scavenger beetles
(incl. Georissidea=Georyssidae,
Helophoridae, Hydrochidae, Spercheidae, Sphaeridiidae)
HELOPHORINAE Leach, 1815
EPIMETOPINAE Zaitzev, 1908
GEORISSINAE Laporte, 1840
HYDROCHINAE C. G. Thomson, 1859
SPERCHEINAE Erichson, 1837
HORELOPHINAE Hansen, 1991
HYDROPHILINAE Latreille, 1802
SPHAERIDIINAE Latreille, 1802
19. SPHAERITIDAE Shuckard, 1839 - False clown beetles
20. SYNTELIIDAE Lewis, 1882
21. HISTERIDAE Gyllenhal, 1808 - Hister or clown beetles
(= Niponiidae)
NIPONIINAE Fowler, 1912
ABRAEINAE MacLeay, 1819
TRYPETICINAE Bickhardt, 1914
TRYPANAEINAE Marseul, 1857
SAPRININAE Blanchard, 1845
DENDROPHILINAE Reitter, 1909
ONTHOPHILINAE MacLeay, 1819
TRIBALINAE Bickhardt, 1914
HISTERINAE Gyllenhal, 1808
HETAERIINAE Marseul, 1857
CHLAMYDOPSINAE Bickhardt, 1914
Superfamily STAPHYLINOIDEA Latreille, 1802
22. HYDRAENIDAE - Minute
moss beetles
(= Limnebiidae)
PROSTHETOPINAE Perkins in Perkins
& Balfour-Browne, 1994
HYDRAENINAE Mulsant, 1844
OCHTHEBIINAE C. G. Thomson,
1859
23. PTILIIDAE Erichson, 1845/Motschulsky, 1845 - Feather-winged beetles
(=Trichopterygidae; incl. Cephaloplectidae,
Limulodidae)
PTILIINAE Erichson, 1845/Motschulsky,
1845
NANOSELLINAE Barber, 1924
CEPHALOPLECTINAE Sharp, 1883
ACROTRICHINAE Reitter, 1909
24. ARGYRTIDAE C. G. Thomson, 1859
(=Silphidae, part)
25. LEIODIDAE Fleming, 1821 - Round fungus beetles
(= Anisotomidae, Liodidae; incl. Camiaridae, Catopidae, Cholevidae, Colonidae,
Leptinidae, Leptodiridae, Platypsyllidae, Sogdiidae)
CAMIARINAE Jeannel, 1911
CATOPOCERINAE Hatch, 1927
LEIODINAE Fleming, 1821
COLONINAE Horn, 1880
CHOLEVINAE Kirby, 1837
PLATYPSYLLINAE Ritsema, 1869
26. SCYDMAENIDAE Leach, 1815 - Ant-like stone beetles
(incl. Anisosphaeridae)
MASTIGINAE Fleming, 1821
SCYDMAENINAE Leach, 1815
27. SILPHIDAE Latreille, 1807- Carrion beetles
SILPHINAE
Latreille, 1807
NICROPHORINAE Kirby, 1837
28. STAPHYLINIDAE Latreille, 1802 - Rove beetles
(incl. Brathinidae, Dasyceridae, Empelidae, Micropeplidae, Oxytelidae,
Oxyporidae, Pselaphidae, Scaphidiidae, etc.)
GLAYPHOLOMATINAE Jeannel, 1962
MICROSILPHINAE Crowson, 1950
OMALIINAE MacLeay, 1825
EMPELINAE Newton & Thayer,
1992
PROTEININAE Erichson, 1839
MICROPEPLINAE Leach, 1815
NEOPHONINAE Fauvel, 1905
DASYCERINAE Reitter, 1887
PROTOPSELAPHINAE Newton &
Thayer, 1995
PSELAPHINAE Latreille, 1802
PHLOEOCHARINAE Erichson, 1839
OLISHAERINAE C. G. Thomson,
1858
TACHYPORINAE MacLeay, 1825
TRICHOPHYINAE C. G. Thomson,
1858
HABROCERINAE Mulsant & Rey,
1877
ALEOCHARINAE Fleming, 1821
TRIGONURINAE Reiche, 1865
APATETICINAE Fauvel, 1895
SCAPHIDIINAE Latreille, 1807
PIESTINAE Erichson, 1839
OSORIINAE Erichson, 1839
OXYTELINAE Fleming, 1821
OXYPORINAE Fleming, 1821
MEGALOPSIDIINAE Leng, 1920
STENINAE MacLeay, 1825
EUAESTHETINAE C. G. Thomson,
1859
SOLIERIINAE Newton & Thayer,
1992
LEPTOTYPHLINAE Fauvel, 1874
PSEUDOPSINAE Ganglbauer, 1895
PAEDERINAE Fleming, 1821
STAPHYLININAE Latreille, 1802
Series SCARABAEIFORMIA Crowson, 1960
Superfamily SCARABAEOIDEA
Latreille, 1802
(= Lamellicornia)
29. LUCANIDAE Latreille, 1804 - Stag beetles
AESALINAE MacLeay, 1819
NICAGINAE LeConte, 1861
SYNDESINAE MacLeay, 1819
LAMPRIMINAE MacLeay, 1819
PENICHROLUCANINAE Arrow, 1950
LUCANINAE Latreille, 1804
30. PASSALIDAE Leach, 1815 - Bess beetles
AULACOCYCLINAE Kaup, 1868
PASSALINAE Leach, 1815
31. TROGIDAE MacLeay, 1819 - Hide beetles
32. GLARESIDAE Semenov-Tian-Shanskij & Medvedev,
1932 - Enigmatic scarab beetles
33. PLEOCOMIDAE LeConte, 1861 - Rain beetles
34. DIPHYLLOSTOMATIDAE Holloway, 1972 - False stag beetles
35. GEOTRUPIDAE Latreille, 1802 - Earth-boring scarab beetles
BOLBOCERATINAE Mulsant, 1842
GEOTRUPINAE Latreille, 1802
LETHRINAE Mulsant & Rey,
1871
36. BELOHINIDAE Paulian, 1959
37. OCHODAEIDAE Mulsant & Rey, 1871 - Sand-loving scarab beetles
OCHODAEINAE Mulsant & Rey,
1871
CHAETOCANTHINAE Scholtz in Scholtz,
et al., 1988
38. CERATOCANTHIDAE Cartwright & Gordon, 1971
- Pill scarab beetles
(= Acanthoceridae)
39. HYBOSORIDAE Erichson, 1847 - Scavanger scarab beetles
40. GLAPHYRIDAE MacLeay, 1819 - Bumble bee scarab beetles
(incl. Lichniidae)
41. SCARABAEIDAE Latreille, 1802 - Scarabs, dung beetles, May/June beetles, chafers
(= Aclopidae, Cetoniidae, etc.)
APHODIINAE Leach, 1815
SCARABAEINAE Latreille, 1802
PACHYPODINAE Erichson, 1840
ORPHNINAE Erichson, 1847
ALLIDIOSTOMATINAE Arrow, 1940
ACLOPINAE Milne-Edwards, 1850
EUCHIRINAE Hope, 1840
PHAENOMERIDINAE Erichson, 1847
MELOLONTHINAE Macleay, 1819/Leach
in Samouelle, 1819
RUTELINAE MacLeay, 1819
DYNASTINAE MacLeay, 1819
CETONIINAE Leach, 1815
Series ELATERIFORMIA
Crowson, 1960
(= Dascilliformia; incl. Eucinetiformia)
Superfamily SCIRTOIDEA Fleming, 1821
42. DECLINIIDAE Nikisky et al., 1994
43. EUCINETIDAE Lacordaire, 1857 - Plate-thigh beetles
(= Cryptomeridae)
44. CLAMBIDAE Fischer, 1821 - Minute beetles
(incl. Calyptomeridae)
CALYPTOMERINAE Crowson, 1955
ACALYPTOMERINAE Crowson, 1979
CLAMBINAE Fischer, 1821
45. SCIRTIDAE Fleming, 1821 - Marsh beetles
(= Cyphonidae, Helodidae, Elodidae)
Superfamily DASCILLOIDEA Guérin-Méneville, 1843
46. DASCILLIDAE Guérin-Méneville, 1843
- Soft-bodied plant beetles
(incl. Karumiidae)
DASCILLINAE Guérin-Méneville,
1843
KARUMIINAE Escalera, 1913
47. RHIPICERIDAE Latreille, 1834 - Cedar beetles or cicada parasite beetles
(= Sandalidae)
Superfamily BUPRESTOIDEA Leach, 1815
48. SCHIZOPODIDAE LeConte, 1861 - Schizopodid or false jewel beetles
49. BUPRESTIDAE Leach, 1815 - Metallic wood-boring or jewel beetles
JULODINAE
Lacordaire, 1857
POLYCESTINAE Lacordaire, 1857
GALBELLINAE Reitter, 1911
CHALCOPHORINAE Lacordaire, 1857
BUPRESTINAE Leach, 1815
AGRILINAE Laporte, 1835
Superfamily BYRROIDEA
Latreille, 1804
(incl. Dryopoidea, Psephenoidea)
50. BYRRHIDAE Latreille, 1804 - Pill or moss beetles
(incl. Syncalyptidae)
BYRRHINAE Latreille, 1804
SYNCALYPTINAE Mulsant &
Rey, 1869
AMPHICYRTINAE LeConte, 1861
51. ELMIDAE Curtis, 1830 - Riffle beetles
(
= Elminthidae, Helminthidae, Elmididae, Limniidae)
LARAINAE LeConte, 1861
ELMINAE Curtis, 1830
52. DRYOPIDAE Billberg, 1820 - Long-toed water beetles
(= Parnidae; incl. Chiloeidae)
53. LUTROCHIDAE Kasap & Crowson, 1975 - Travertine beetles
54. LIMNICHIDAE Erichson, 1846 - Minute marsh-loving beetles
HYPHALINAE Britton, 1971
LIMNICHINAE Erichson, 1846
CEPHALOBYRRHINAE Champion, 1925
THAUMASTODINAE Champion, 1924
55. HETEROCERIDAE MacLeay, 1825 - Variegated mud-loving beetles
ELYTHOMERINAE Pacheco, 1964
HETEROCERINAE MacLeay, 1825
56. PSEPHENIDAE Lacordaire, 1854 - Water penny beetles
(incl. Eubriidae, Psephenoididae)
EUBRIINAE Lacordaire, 1857
PSEPHENOIDINAE Hinton, 1939
EUBRIANACINAE Jacobson, 1913
PSEPHENINAE Lacordaire, 1854
57. CNEOGLOSSIDAE Champion, 1897
58. PTILODACTYLIDAE Laporte, 1836 - Toed-winged beetles
ANCHYTARSINAE Champion, 1897
CLADOTOMINAE Pic, 1914
APLOGLOSSINAE Champion, 1897
ARAEOPIDIINAE Lawrence, 1991
PTILODACTYLINAE Laporte, 1836
59. CHELONARIIDAE Blanchard, 1845
60. EULICHADIDAE Crowson, 1973 - Forest stream beetles
(= Lichadidae)
61. CALLIRHIPIDAE Emden, 1924
Superfamily ELATEROIDEA
Leach, 1815
(incl. Cantharoidea, Artematopodoidea)
62. ARTEMATOPODIDAE Lacordaire, 1857
(= Artematopidae; incl. Eurypogonidae)
ELECTRIBIINAE Crowson, 1975
ARTEMATOPODINAE Lacordaire,
1857
63. BRACHYPSECTRIDAE LeConte & Horn, 1883 - Texas beetles
64. CEROPHYTIDAE Latreille, 1834 - Rare click beetles
65. EUCNEMIDAE Eschscholtz, 1829 - False click beetles
(= Melasidae; incl. Perothopidae,
Phylloceridae)
PEROTHOPINAE Lacordaire, 1857
PHYLLOCERINAE Reitter, 1905
PSEUDOMENINAE Muona, 1993
PALAEOXENINAE Muona, 1993
PHLEOGONINAE Muona, 1993
MELASINAE Fleming, 1821
EUCNEMINAE Eschscholtz, 1829
MACRAULACINAE Fleutiaux, 1923
66. THROSCIDAE Laporte, 1840
(= Trixagidae)
67. ELATERIDAE Leach, 1815 - Click beetles
(incl. Cebrionidae, Dicronychidae, Lissomidae)
CEBRIONINAE Latreille, 1802
TETRALOBINAE Laporte, 1840
THYLACOSTERNINAE Fleutiaux,
1920
LISSOMINAE Laporte, 1835
SEMIOTINAE Jacobson, 1913
PITYOBIINAE Hyslop, 1917
OXYNOPTERINAE Candèze,
1857
AGRYPNINAE Candèze, 1857/Lacordaire,
1857
DENTICOLLINAE Stein & Weise,
1877
NEGASTRIINAE Nakane & Kishii,
1956
DIMINAE Candèze, 1863
ELATERINAE Leach, 1815
CARDIOPHORINAE Candèze,
1859
HEMIOPINAE Fleutiaux, 1941
PHYSODACTYLINAE Lacordaire,
1857
incertae sedis
EUDICHRONYCHINAE Girard, 1971
ANISCHIINAE Fleutiaux, 1936
SUBPROTELATERINAE Fleutiaux,
1920
68. PLASTOCERIDAE Crowson, 1972
69. DRILIDAE Blanchard, 1845
70. OMALISIDAE Lacordaire, 1857
(= Omalysidae, Homalisidae)
71. LYCIDAE Laporte, 1836 - Net-winged beetles
LYCINAE Laporte, 1836
LEPTOLYCINAE Leng & Mutchler,
1922
ATELIINAE Kleine, 1928
METRIORRHYNCHINAE Kleine, 1928
EROTINAE LeConte, 1881
CALOCHROMINAE Lacordaire, 1857
72. TELEGEUSIDAE Leng, 1920 - Long-lipped beetles
73. PHENGODIDAE LeConte, 1861 - Glow-worms
RHAGOPHTHALMINAE Olivier, 1907
PHENOGODINAE LeConte, 1861
74. LAMPYRIDAE Latreille, 1817 - Fireflies, Lightingbugs, Glowworms
PTEROTINAE LeConte, 1861
OTOTRETADRILINAE Crowson, 1972
CYPHONOCERINAE Crowson, 1972
OTOTRETINAE McDermott, 1964
AMYDETINAE Olivier, 1907
LAMPYRINAE Latreille, 1817
LUCIOLINAE Lacordaire, 1857
PHOTURINAE Lacordaire, 1857
75. OMETHIDAE LeConte, 1861 - False solider beetles
DRILONIINAE Crowson, 1972
MATHETEINAE LeConte, 1881
OMETHINAE LeConte, 1861
76. CANTHARIDAE Imhoff, 1856 - Soldier beetles
(= Telephoriade;
incl. Chauliognathidae)
CANTHARINAE Imhoff, 1856
SILINAE Mulsant, 1862
DYSMORPHOCERINAE Brancucci,
1980
MALTHININAE Kiesenwetter, 1852
CHAULIOGNATHINAE LeConte, 1861
ELATERIFORMIA Incertae Sedis
77. PODOCEPHALIDAE Pic, 1930
78. RHINORHIPIDAE Lawrence, 1988
Series BOSTRICHIOFORMIA Forbes, 1926
Superfamily DERODONTOIDEA LeConte, 1861
79. DERODONTIDAE LeConte, 1861 - Tooth-neck fungus beetles
(incl. Laricobiidae,
Peltasticidae)
PELTASTICINAE LeConte, 1861
DERODONTINAE LeConte, 1861
LARICOBIINAE Mulsant & Rey,
1863-64
Superfamily BOSTRICHOIDEA
Latreille, 1802
(inc. Dermestoidea)
80. NOSODENDRIDAE Erichson, 1846
81. DERMESTIDAE Latreiile, 1804 - Skin beetles
(incl. Thorictidae, Thylodriidae)
DERMESTINAE Latreille, 1804
MARIOUTINAE Jacobson, 1913
THORICTINAE Agassiz, 1846
ORPHILINAE LeConte, 1861
TRINODINAE Casey, 1900
THYLODRIINAE Semenov-Tian-Shanskij,
1913
ATTAGENINAE Laporte, 1840
MEGATOMINAE Leach, 1815
82. ENDECATOMIDAE LeConte, 1861
83. BOSTRICHIDAE Latreille, 1802 - Powder-post beetles
(= Bostrychidae; incl. Lyctidae, Psoidae)
DYSIDINAE Lesne, 1921
POLYCAONINAE Lesne, 1896
BOSTRICHINAE Latreille, 1802
PSOINAE Blanchard, 1851
DINODERINAE C. G. Thomson, 1963
LYCTINAE Billberg, 1820
EUDERIINAE Lesne, 1934
84. ANOBIIDAE Fleming, 1821 - Death watch and spider beetles
(incl. Ectrephidae, Gnostidae, Ptinidae)
EUCRADINAE LeConte, 1861
PTININAE Latreille, 1802
DRYOPHILINAE LeConte, 1861
ERNOBIINAE Pic, 1912
ANOBIINAE Fleming, 1821
PTILININAE Shuckard, 1840
ALVARENGANIELLINAE Viana &
Martinez,
XYLETININAE GIstel, 1856
DORCATOMINAE C. G. Thomson,
1859
MESOCOELOPODINAE Mulsant &
Rey, 1864
BOSTRICHIOFORMIA Incertae Sedis
85. JACOBSONIIDAE Heller, 1926
(= Sarothriidae)
Series CUCUJIFORMIA Lameere, 1938
Superfamily LYMEXYLONIODEA Fleming, 1821
86. LYMEXYLIDAE Fleming, 1821 - Ship-timber beetles
(= Lymexylonidae;
incl. Atractoceridae)
HYLECOETINAE
Gistel, 1856
LYMEXYLINAE Fleming, 1821
MELITTOMMATINAE Wheeler, 1986
Superfamily CLEROIDEA Latrielle, 1802
87. PHLOIOPHILIDAE Kiesenwetter, 1863
(= Phloeophilidae)
88. TROGOSSITIDAE Latreille, 1802 - Bark-gnawing beetles
(incl. Lophocateridae, Ostomidae,
Peltidae, Temnochilidae)
PROTOPELTINAE Crowson, 1966
LARINOTINAE Slipinski, 1992
PETINAE Kirby, 1837
RENTONIINAE Crowson, 1966
DECAMERINAE Crowson, 1964
LOPHOCATERINAE Crowson, 1964
CALITINAE Reitter, 1922
EGOLIINAE Lacordaire, 1854
TROGOSSITINAE Latreille, 1802
89. CHAETOSOMATIDAE Crowson, 1952
90. CLERIDAE Latreille, 1802 - Checkered beetles
(incl. Corynetidae, Korynetidae)
THANEROCLERINAE Chapin, 1924
TILLINAE Leach, 1815
HYDNOCERINAE Spinola, 1844
CLERINAE Latreille, 1802
EPIPHLOEINAE Kuwert, 1893
ENOPLIINAE Gistel, 1856
TARSOSTENINAE Jacquelin du Val,
1860
KORYNETINAE Laporte, 1836
91. ACANTHOCNEMIDAE Crowson, 1964
92. PHYCOSECIDAE Crowson, 1952
93. PRIONOCERIDAE Lacordaire, 1857
94. MELYRIDAE Leach, 1815 - Soft-winged flower beetles
(incl. Dasytidae, Malachiidae, Rhadalidae,
Carphuridae)
MELYRINAE
Leach, 1815
RHADALINAE LeConte, 1861
GIETELLINAE Constantin &
Menier, 1987
DASYTINAE Laporte, 1840
MALACHINAE Fleming, 1821
Superfamily CUCUJOIDEA
Latreille, 1802 (= Clavicornia)
95. PROTOCUCUJIDAE Crowson, 1954
96. SPHINDIDAE Jacquelin du Val, 1860 - Dry-fungus beetles
(incl. Aspidiphroidae)
PROTOSPHINDINAE Sen Gupta &
Crowson, 1979
ODONTOSPHINDINAE
Sen Gupta & Crowson, 1979
SPHINDIPHORINAE
Sen Gupta & Crowson, 1979
SPHINDINAE
Jacquelin du Val, 1860
97. BRACHYPTERIDAE Erichson, 1845 - Short-winged flower beetles
(= Cateretidae, Kateretidae)
98. NITIDULIDAE Latreille, 1802 - Sap beetles
(incl. Cybocephalidae)
CALONECRINAE Kirejtshuk, 1982
CARPOPHILINAE Erichson, 1842
MELIGETHINAE C. G. Thomson,
1859
NITIDULINAE Latreille, 1802
CILLAEINAE Kirejtshuk &
Audisio, in Kirejtshuk, 1986
CRYPTARCHINAE C. G. Thomson,
1859
CYBOCEPHALINAE
Jacquelin du Val, 1858
99. SMICRIPIDAE Horn, 1879 -
Palmetto beetles
100. MONOTOMIDAE Laporte, 1840 -
Root-eating beetles
(incl. Rhizophagidae)
RHIZOPHAGINAE Redtenbacher, 1845
MONOTOMINAE Laporte, 1840
101. BOGANIIDAE Sen Gupta & Crowson, 1966
PARACUCUJINAE Endrody-Younga
& Crowson, 1986
BOGANIINAE Sen Gupta & Crowson,
1966
102. HELOTIDAE Reitter, 1876/Chapuis, 1876
103. PHLOEOSTICHIDAE Reitter, 1911
PHLOEOSTICHINAE Reitter, 1911
HYMAEINAE
Sen Gupta & Crowson, 1969
AGAPYTHINAE Sen Gupta & Crowson,
1969
MYRABOLIINAE
Lawrence & Britton, 1991
PRIASILPHINAE Crowson, 1973
TASMOSALPINGINAE Lawrence &
Britton, 1991
104. SILVANIDAE Kirby, 1837 - Silvanid
flat bark beetles
BRONTINAE Erichson, 1845/Blanchard,
1845
SILVANINAE Kirby, 1837
105. PASSANDRIDAE Erichson, 1845/Blanchard, 1845
(= Catogenidae)
106. CUCUJIDAE Latreille, 1802 - Flat bark beetles
107. LAEMOPHLOEIDAE Ganglbauer, 1899 - Lined flat bark beetles
108. PROPALTICIDAE Crowson, 1952
109. PHALACRIDAE Leach, 1815 - Shining
flower beetles
(incl.
Phaenocephalidae)
PHAENOCEPHALINAE Matthews, 1899
PHALACRINAE Leach, 1815
110. HOBARTIIDAE Sen Gupta & Crowson, 1966
111. CAVOGNATHIDAE Sen Gupta & Crowson, 1966
112. CRYPTOPHAGIDAE Kirby, 1837 - Silken
fungus beetles
(incl.
Catopochrotidae, Hypocopridae)
HYPOCOPRINAE Reitter, 1879
CRYPTOPHAGINAE Kirby, 1837
ATOMARIINAE LeConte, 1861
113. LAMINGTONIDAE
Sen Gupta & Crowson, 1966
114. LANGURIIDAE Crotch, 1873 - Lizard
beetles
(incl. Cryptophilidae)
XENOSCELINAE Ganglbauer, 1899
SETARIOLINAE Crowson, 1952
LANGURIINAE Crotch, 1873
CRYPTOPHILINAE Casey, 1900
TORAMINAE Sen Gupta, 1967
115. EROTYLIDAE Latreille, 1802 -
Pleasing fungus beetles
(incl. Dacnidae)
DACNINAE Gistel, 1856
MEGALODACNINAE Sen Gupta, 1969
ENCAUSTINAE Crotch, 1876/Chapuis,
1876
TRITOMINAE Curtis, 1834
EROTYLINAE Latreille, 1802
116. BYTURIDAE - Fruitworms
PLATYDASCILLINAE Pic, 1914
BYTURIDAE Jacquelin du Val,
1858
117. BIPHYLLIDAE LeConte, 1861 -
False skin beetles
(= Diphyllidae)
118. BOTHRIDERIDAE Erichson, 1845
(incl. Anommatidae)
TEREDINAE Seidlitz, 1888
XYLARIOPHILINAE Pal & Lawrence,
1986
ANOMMATINAE Ganglbauer, 1899
BOTHRIDERINAE Erichson, 1845
119. CERYLONIDAE Billberg, 1820 - Minute bark beetles
(= Cerylidae; incl. Aculagnathidae, Dolosidae, Euxestidae, Murmidiidae)
EUXESTINAE Grouvelle, 1908
LOEBLIORYLONINAE
Slipinski, 1990
OSTOMOPSINAE Sen
Gupta & Crowson, 1973
MURMIDIINAE Jacqueline
du Val, 1858
CERYLONINAE Billberg,
1820
120. ALEXIIDAE Imhoff, 1856
(= Sphaerosomatidae, Sphaerosomidae)
121. DISCOLOMATIDAE Horn, 1878
(= Discolomidae, Notiophygidae)
NOTIOPHYGINAE Jacobson, 1915
DISCOLOMATINAE Horn, 1878
APHANOCEPHALINAE Grouvelle,
1912
CEPHALOPHANINAE John, 1954
PONDONATINAE John, 1954
122. ENDOMYCHIDAE Leach, 1815 -
Handsome fungus beetles
(incl. Merophysiidae, Mycetaeidae)
MEROPHYSIINAE Seidlitz, 1872
HOLOPARAMECINAE Seidlitz,
1888
LEIESTINAE C. G.
Thomson, 1863
EUPSILOBIINAE Casey,
1895
ENDOMYCHINAE Leach,
1815
EPIPOCINAE Gorham,
1873
LYCOPERDININAE Redtenbacher,
1844
MYCETAEINAE Jacquelin
du Val, 1857
ANAMORPHINAE Strohecker,
1953
PLEGANOPHORINAE Jacquelin
du Val, 1858
XENOMYCETINAE Strohecker
in Arnett, 1962
123. COCCINELLIDAE Latreille, 1807 - Ladybird beetles
(incl. Epilachnidae)
STICHOLOTIDINAE Weise, 1901
COCCIDULINAE Mulsant, 1846
SCYMNINAE Mulsant, 1846
CHILOCORINAE Mulsant, 1846
COCCINELLINAE Latreille,
1807
EPILACHNINAE Mulsant, 1846
124. CORYLOPHIDAE LeConte, 1852 - Minute fungus beetles
(= Orthoperidae)
PELTINODINAE Paulian, 1950
CORYLOPHINAE LeConte,
1852
SERICODERINAE Matthews, 1888
PARMULINAE Poey,
1854
125. LATRIDIIDAE Erichson, 1842 - Minute brown scavenger beetles
LATRIDIINAE Erichson, 1842
CORTICARIINAE Curtis, 1829
Superfamily TENEBRIONOIDEA
Latreille, 1802
(= Heteromera)
126. MYCETOPHAGIDAE Leach, 1815 - Hairy fungus beetles
ESARCINAE Reitter, 1882
MYCETOPHAGINAE Leach, 1815
BERGININAE Leng, 1920
127. ARCHEOCRYPTICIDAE Kaszab, 1964
128. PTEROGENIIDAE Crowson, 1953
129. CIIDAE Leach in Samouelle, 1819 - Minute tree-fungus beetles
(= Cisidae )
SPHINDOCIINAE Lawrence, 1974
CIINAE Leach in Samouelle,
1819
130. TETRATOMIDAE Billberg, 1820 -
Polypore fungus beetles
PISENINAE Miyatke, 1960
TETRATOMINAE Billberg, 1820
PENTHINAE Lacordaire, 1859
131. MELANDRYIDAE Leach, 1815 -
False farkling beetles
(= Serropalpidae)
HALLOMENINAE Mulsant, 1856/Gistel,
1856
EUSTROPHINAE Gistel, 1856
MELANDRYINAE Leach, 1815
OSPHYINAE Mulsant, 1856
132. MORDELLIDAE Latreille, 1802 - Tumbling flower beetles
CTENIDINNAE Franciscolo, 1851
MORDELLINAE Latreille, 1802
133. RHIPIPHORIDAE Gemminger & Harold, 1870
PELECOTOMINAE Seidlitz, 1875
MICHOLAEMINAE Viana, 1971
PTILOPHORINAE
Gerstaecker, 1855
HEMIRHIPIDIINAE Heller, 1921
RHIPIDIINAE Gerstaecker, 1855
RHIPIPHORINAE Germminger &
Harold, 1870
134. COLYDIIDAE Erichson, 1842 -
Cylindrical bark beetles
(incl. Adimeridae, Monoedidae)
PYCNOMERINAE Erichson, 1845
COLYDIINAE Erichson,
1842
135. MONOMMATIDAE Blanchard, 1845
(= Monommidae)
136. ZOPHERIDAE Solier, 1834 - Ironclad
beetles
(incl. Merycidae, part)
ZOPHERINAE Solier, 1834
USECHINAE Horn, 1867
137. ULODIDAE Pascoe, 1869
(incl. Merycidae, part)
138. PERIMYLOPIDAE St. George, 1939
139. CHALCODRYIDAE Watt, 1974
140. TRACHELOSTENIDAE Lacordaire, 1859
141. TENEBRIONIDAE Latreille, 1802 - Darkling beetles
(incl. Alleculidae, Cossyphodidae, Lagriidae, Nilionidae, Petriidae,
Physopaussidae, Tentyriidae)
LAGRIINAE Latreille, 1825
PHRENAPATINAE Solier,
1834
ZOLODININAE Watt,
1974
PIMELIINAE Latreille,
1802
TENEBRIONINAE Latreille,
1802
ALLECULINAE Laporte,
1840
DAIPERINAE Latreille,
1802
COELOMETOPINAE Lacordaire,
1859/Schaum, 1859
incertae
sedis
COSSYPHDINAE Wamann,
1899
142. PROSTOMIDAE C. G. Thomson, 1859 - Jugular-horned beetles
143. SYNCHROIDAE Lacordaire, 1859 - Synchroa bark beetles
144. OEDEMERIDAE Latreille, 1810 - False blister beetles
CALOPODINAE Costa, 1852
OEDEMERINAE Latreille, 1810
145. STENOTRACHELIDAE C. G. Thomson, 1859 - False longhorn beetles
(= Cephaloidae)
STENOTRACHELINAE C. G.
Thomson, 1859
NEMATOPLINAE LeConte, 1852
CEPHALOINAE LeConte, 1852
incertae sedis
STOLIINAE Nikitsky, 1985
146. MELOIDAE Gyllenhal, 1810 - Blister beetles
(incl. Tetraonychidae)
ELETICINAE Wellman, 1910
MELOINAE Gyllenhal,
1810
NEMOGNATHINAE Laporte,
1840
147. MYCTERIDAE Blanchard, 1845 - Palm and flower beetles
(incl. Hemipeplidae)
MYCTERINAE Blanchard, 1845
LACCONOTINAE LeConte, 1862
HEMIPEPLINAE Lacordaire,
1854
148. BORIDAE
C. G. Thomson, 1859 - Conifer bark beetles
BORINAE C. G. Thomson, 1859
SYNERCTICINAE Lawrence &
Pollock, 1994
149. TRICTENOTOMIDAE Blanchard, 1845
150. PYTHIDAE Solier, 1834 -
Dead log beetles
151. PYROCHROIDAE Latreille, 1807 - Fire-colored beetles
(incl. Conontidae, Pedilidae, Pilipalpidae)
TYDESSINAE Nikitshy, 1986
PILIPALPINAE Abdullah, 1964
PEDILINAE Lacordaire, 1859
PYROCHROINAE Latreille, 1807
incertae sedis
AGNATHINAE Lacordaire, 1859
152. SALPINGIDAE Leach, 1815 - Narrow-waisted bark beetles
(incl. Aegialitidae, Dacoderidae, Elacatidae, Eurystethidae, Inopeplidae,
Othiniidae, Tretothoracidae)
OTHNIINAE LeConte, 1861
PROSTOMINIINAE Grouvelle, 1914
AGLENINAE Horn, 1878
INOPEPLINAE Grouvelle, 1908
SALPINGINAE Leach, 1815
AEGIALITINAE LeConte, 1862
DACODERINAE LeConte, 1862
153. ANTHICIDAE Latreille, 1819 - Ant-like flower beetles
EURYGENIINAE LeConte, 1862
LAGRIOIDINAE Abdullah &
Abdullah, 1968
AFREMINAE Levey, 1985
MACRATRIINAE LeConte, 1862
STEROPINAE Jacquelin du Val,
1863
ISCHALIINAE Blair, 1920
COPOBAENINAE Abdullah, 1969
LEMODINAE Lawrence & Britton,
1991
TOMODERINAE Bonadona, 1961
ANTHICINAE Latreille, 1819
154. ADERIDAE Winkler, 1927 - Ant-like leaf beetles
(= Euglenidae, Euglenesidae, Hylophilidae, Xylophilidae)
155. SCRAPTIIDAE Mulsant, 1856/Gistel, 1856 - False flower beetles
(incl. Anaspididae)
SCRAPTIINAE Mulsant, 1856/Gistel, 1856
ANASPIDINAE Mulsant,
1856
Superfamily CHRYSOMELOIDEA Latreille, 1802
156. CERAMBYCIDAE Latreille, 1802 - Longhorn beetles
(incl. Anoplodermatidae, Disteniidae,
Hypocephalidae, Oxypeltidae, Paradridae, Spondylidae, Vesperidae, etc.)
VESPERINAE Mulsant, 1839
OXYPELTINAE Lacordaire, 1869
DISTENIINAE J. Thomson, 1860
ANOPLODERMATINAE Guérin-Méneville,
1840
PHILINAE J. Thomson, 1860
PARANDRINAE Blanchard, 1845
PRIONINAE Latreille, 1802
SPONDYLIDINAE Audinet-Serville,
1832
APATOPHYSEINAE Lacordaire, 1869
NECYDALINAE Latreille, 1825
LEPTURINAE Latreille, 1802
CERAMBYCINAE Latreille, 1802
LAMIINAE Latreille, 1825
157. MEGALOPODIDAE Latreille, 1802
PALOPHAGINAE Kuschel & May, 1990
ZEUGOPHROINAE Böving &
Craighead, 1931
MEGALOPODINAE Latreille, 1802
158. ORSODACNIDAE C. G. Thomson, 1859
ORSODACNINAE C. G. Thomson, 1859
AULACOSCELIDINAE Chapuis, 1874
159. CHRYSOMELIDAE Latreille, 1802 - Leaf beetles
(incl. Bruchidae, Sagridae, etc.)
SAGRINAE Leach, 1815
BRUCHINAE Latreille, 1802
DONACIINAE Kirby, 1837
CRIOCERINAE Latreille, 1804
HISPINAE Gyllenhal, 1813
CHRYSOMELINAE Latreille, 1802
GALERUCINAE Latreille, 1802
LAMPROSOMATINAE Lacordaire,
1848
CRYPTOCEPHALINAE Gyllenhal,
1813
EUMOLPINAE Hope, 1840
Superfamily CURCULIONOIDEA
Latreille, 1802
(= Rhynchophora)
160. NEMONYCHIDAE Bedel, 1882 -
Pine flower snout beetles
(= Rhinomaceridae)
NEMONYCHINAE Bedel, 1882
RHINORHYNCHINAE Voss,
1922
DOYDIRHYNCHINAE Pierce,
1916
161. ANTHRIBIDAE Billberg, 1820 - Fungus weevils
(= Platystomidae, Platostomatidae, Platyrhinidae; incl. Bruchelidae,
Urodontidae)
URODONATINAE C. G. Thomson, 1859
ANTHRIBINAE Billberg,
1820
CHORAGINAE Kirby,
1819
162. BELIDAE Schönherr, 1826 - Primitive weevils
BELINAE
Schönherr, 1826
OXYCORYNIDAE
Schönherr, 1840
AGLYCYDERINAE Wollaston, 1864
163. ATTELABIDAE Billberg, 1820 - Leaf Rolling weevils
(incl. Apoderidae, Pterocolidae, Rhynchitidae)
RHYNCHITINAE Gistel, 1856
ATTELABINAE Billberg,
1820
164. BRENTIDAE Billberg, 1820
- Straight snouted weevils
(= Brenthidae; incl. Apionidae,
Cyladidae, Eurhynchidae)
EURHYNCHINAE Lacordaire, 1863
CYLADINAE Schönherr, 1823
BRENTINAE Billberg, 1820
ANTLIARHININAE Schönherr, 1823
NANOPHYINAE Gistel, 1856
APIONINAE Schönherr,
1823
165. CARIDAE Thompson, 1992
166. ITHYCERIDAE Schönherr, 1823 - New York weevil
167. CURCULIONIDAE Latreille, 1802 - Weevils or snout beetles
(incl. Calendridae, Cossonidae, Platypodidae, Rhyncophoridae, Scolytidae,
etc.)
BRACHYCERINAE Billberg, 1820
CURCULIONINAE Latreille, 1802
DRYOPHTHORINAE Schönherr,
1825
COSSONINAE Schönherr, 1825
SCOLYTINAE Latreille, 1804
PLATYPODINAE Shuckard, 1840
